Atlassian has posted the following relevant information regarding Atlassian Marketplace apps/add-ons and development licenses of JIRA:
Which points to:
Licensing and Paid via Atlassian Listings
Go to the section titled, "Can customers use developer licenses for my add-on?", which has instructions on how to get a developer license for apps/add-ons.
A developer license can be generated (free of charge) once the plugin has been purchased and can only be installed on a development licensed version of JIRA.
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